You and I both know people or dogs that we don't consider
particularly sophisticated.
We sometimes refer to them as "simple" or "Real Housewives."
But when it comes to truly simple animals,
we shouldn't underestimate them.
Because the animal phyla that we describe as being the least
complex actually offer us a vivid way of understanding how animals
are structured, and also how they evolved.
Simple doesn't always mean dumb.
Unlike those dullards that we've all met in our lives,
animals aren't considered simple because they apparently take things
for "granite" or they think that reality TV is...reality.
Their simplicity has to do with their tissue complexity.
As you know, almost all animals' cells are organized into tissues
that perform specialized functions.
The more different kinds of specialized cells an animal has,
the more complex it is, and this complexity is determined
in the embryonic phase.
As embryos, most animals either form two layers of early tissue,
called germ layers, or they form three.
By exploring the very simplest phyla, from animals with no
layers at all, aka sponges, to the most basic of three-layer animals,
like mollusks you can see how a not-totally-amazing-sounding change
in tissue results in truly fundamental and amazing changes.
So the places in the animal family tree where these transitions
take place, from no layers to two layers,
and from two layers to three, are some of the most important
benchmarks in animal evolution.
Let's start with the very simplest of animals,
in the phylum Porifera: the sponges!
They diverged from protists probably 600 million years ago and
not a whole lot has changed for them since then.
If you've been paying attention, you've noticed by now that almost
nothing that applies to other animals applies to sponges.
That's because they're so freaking simple.
They can't move; they just hang out and filter water for food
like bacteria, while some host photosynthesizing microbes
and mooch off them.
More important, sponge embryos don't have any layers,
they just have cells.
This means that sponges don't have specialized tissues or organs.
And their cells can take different forms.
Some have flagella to force water into the sponge;
some are more amoeba-like and wander around
distributing nutrients.
But these cells can transform into whatever type of cell
the sponge needs.
For this reason, some scientists argue that sponges aren't even
animals at all, they're actually colonies of cells that depend on
each other to function.
But for our purposes, mainly because they're multicellular,
eukaryotic organisms that can't make their own food,
they still count.
And they've managed to diversify into nearly
10,000 different species, so good for them.
Things get more interesting with Cnidaria, which include jellies,
sea anemones, corals, and hydras.
They got a couple of sweet evolutionary breaks that made them animals that you do NOT want
to mess with.
The first and most important break is that they develop
two germ layers. You'll remember these layers
are called the endoderm, or the "inside" -derm,
and the ectoderm, the "outside" -derm,
and they form a tube that allows an animal to ingest,
digest and get rid of stuff.
This makes Cnidaria among the oldest living descendants of the
world's first diploblast, which is the common ancestor
of all "true animals."
But still, jellies and anemones and other cnidarians have only
one hole that serves as both mouth and anus,
and they don't have any organs.
So, still pretty simple.
Their second evolutionary break is in their ectoderm,
which contains stinging cells called cnidocysts.
Think Portuguese Man o' War, I once stepped on a dead one,
it was dead, LONG dead, and I wanted someone to
cut my foot off it hurt so much.
So now we've got two-layer animals swimming around,
able to move and eat and poop and defend themselves.
The animal kingdom is just one evolutionary breakthrough away
from a huge, like,
explosion!
And we can see the evidence of this breakthrough in Platyhelminthes,
the phylum of soft, unsegmented worms that includes flatworms,
planaria, tapeworms, and flukes.
Not super-handsome, but these guys are a big deal,
because they're the oldest existing phylum that is triploblastic,
or has three germ layers.
So in addition to an endoderm and ectoderm,
their embryos form a mesoderm.
I know it sounds like just another piece of toast and turkey
on a club sandwich, but this development changes everything.
Platyhelminthes are themselves pretty simple,
but a couple of phyla up the ranks,
this new layer allows animals to form true organ systems:
the ectoderm forming the brain, nervous system and skin;
the mesoderm forming muscles, bones, cartilage, the heart,
blood and other very useful stuff;
and the endoderm forming the digestive and respiratory systems.
And this kind of complexity is only possible because of
one of the mesoderm's key features, the coelom,
a fluid-filled cavity that stores and protects the major organs.
It allows the internal organs to move independent of the body wall,
and the fluid can provide some shock resistance.
Coeloms are where all the action happens when it comes
to organ systems, but not all triploblasts have them.
From here on, we can assess the complexity of an animal
by whether it has a coelom or not, and if so, how complete it is.
For instance, because they're the simplest of the triploblasts,
Platyhelminthes have their mouths and buttholes on
opposite ends of their bodies, which is awesome for them!
But they're acoelomates, they don't have a coelom,
which tells us they're still on the
shallow end of the pool, complexity-wise.
To give you an idea of how simple, you can cut
a Platyhelminthes in half, and both of the
pieces will happily continue on with their wormy business.
That, my friends, is simplicity.
Now, you probably haven't forgotten that I mentioned
an explosion a minute ago.
Well, I'm not going to taunt you with
talk of explosions without giving you one.
[BIOLO-GRAPHY]
The Cambrian Explosion!
Not long after germ layers became a thing, say 535 million years ago,
life on earth was undergoing some pretty terrific
and rapid innovations.
Over about 10 or 12 million years, about half
of the animal phyla that exist today started to appear.
It remains the most biologically productive period in history.
Think of the most exciting, vibrant, creative,
dangerous experience and then invite all of Kingdom Animalia
to the party.
Like Burning Man, ComicCon, and Coachella all at once.
This is when animals started to look and behave
as we know them today.
Before the Cambrian, most of the big animals
were slow and soft-bodied and ate algae or scavenged.
But this explosion of diversity brought all kinds
of new adaptations, including predatory ones,
like claws, and defensive ones like spikes and armored plates.
Shells and mineral skeletons made their first appearances.
In fact, the adaptations were so many and so abrupt that
in the 1800s the abundance of fossils from this period
was used to argue against evolution.
Scientists offer a lot of different theories about
what caused this explosion.
It was probably a combination of a few of these things.
For one, oxygen levels became very high in Cambrian seas,
which allowed for larger bodies and higher metabolisms.
It's also thought that ocean chemistry changed,
with more minerals becoming available for the production
of shells and skeletons.
And of course, with more diversity comes more competition
and predation, which drove selective pressures
on animals to become either better at hunting
or better at defending themselves.
It's pretty near the top of my list of places I want to go
once I put the finishing touches on my time machine.
But for now, we still have many modern animal phyla
to remind us of this time of crazy awesomeness.
So flukes are cool and all, but things start to get
more complex with another phylum of mostly nasty parasites,
Nematoda, unsegmented roundworms.
These guys are pseudocoelomates, meaning they have
an incomplete body cavity.
Unlike a true coelomate, whose body cavity is
contained within the mesoderm,
pseudocoelomates sort of improvise one
between the mesoderm and the endoderm.
The vast majority of nematodes live in soil,
where they eat bacteria or fungus or parasitize plant roots.
But humans host at least 50 nematode species,
including hookworms, which burrow into our intestines and treat us
like some kind of food court.
But most nematodes are very very small:
a single teaspoon of forest soil can have several hundred in it!
Rotifera, meanwhile, are tiny filter-feeding animals
that live mostly in fresh or salt water, though some of them
can live in damp soil.
They're also pseudocoelomates like nematodes, and although
they are way smaller than most flatworms,
a big honkin' rotifer is like 2 millimeters long,
they're anatomically more complex, as they have a stomach,
jaws and a little tiny anus.
My favorite fun fact about Rotifera is that many
of its species are known to exist entirely of females,
and they reproduce through unfertilized eggs.
Fossils of rotifers have been found as old as 35 million years,
and in many cases, there's not a dude to be found.
You go girls!
Okay, so now for the big dogs: the phylum Mollusca.
Molluscs might be kind of simple, but they're amazing
and some of them are incredibly smart.
They take four different basic forms:
chitons, snails, bivalves, and octopi and squid.
Now, I realize it can be hard to see how an oyster
and an octopus might be related, but molluscs have
some important similarities:
They all have a visceral mass, which is a true coelom
a body cavity completely within the mesoderm that
contains most of the internal organs.
They also have a big, muscular foot which takes different
forms in each class of mollusc.
They have a mantle, which in some molluscs makes
a shell and in others just covers the visceral mass,
And finally, all molluscs except bivalves have a radula,
or a rasping organ on their mouths that they use to scrape up food.
So, chitons, are these headless marine animals,
covered with a plated shell on one side, and they use
their foot to move around on rocks, scraping off algae
with their radula.
You know about bivalves. They have shells that are
divided into two hinged halves, like clams and scallops.
They're filter feeders, so they trap particles of food
in the mucus that covers their gills.
Snails and slugs are gastropods.
One thing that sets them apart is a process called torsion,
in which the visceral mass twists to the side during
embryonic development, so that by the end of it,
its anus is basically right above it's head.
Most gastropods also have a single, spiraled shell
and most use their radula to graze on algae and plants.
And last, but certainly not least, we have the cephalopods,
which are the kings of the Molluscs, as far as I'm concerned.
Cephalopods include octopi and squid, and they are obviously
a lot different from other molluscs.
For starters, they have tentacles that they use to grab their prey,
which they then bite with their beaks
and immobilize with poisonous saliva.
And the foot of a cephalopod has been modified
into a really powerful muscle that shoots out water
to help it move and steer through the water.
But probably the coolest thing about cephalopods
is how smart they are.
While a typical mollusk might have 20,000 neurons,
an octopus has half a billion neurons.
If you just do a YouTube search for octopus,
you'll find all kinds of videos of them opening jars
and stealing peoples' video cameras.
They're like freaking ocean ninjas.
Cephalopods got skillz.
So remember, simple doesn't equal dumb, there's a lot
to learn from our less-developed cousins.
Next time we'll talk about even more complex animals
and what we have to learn from them.
Until then:
Thank you for watching Crash Course Biology,
If you want to review anything that we discussed in this video
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