An omnivore () is an animal that has the ability to eat and survive on both plant and animal
matter.
Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates,
protein, fat, and fiber, and metabolize the nutrients and energy of the sources absorbed.
Often, they have the ability to incorporate food sources such as algae, fungi, and bacteria
into their diet.Omnivores come from diverse backgrounds that often independently evolved
sophisticated consumption capabilities.
For instance, dogs evolved from primarily carnivorous organisms (Carnivora) while pigs
evolved from primarily herbivorous organisms (Artiodactyla).
What this means is that physical characteristics are often not reliable indicators of whether
an animal has the ability to obtain energy and nutrients from both plant and animal matter.
Owing to the wide range of entirely unrelated organisms independently evolving the capability
to obtain energy and nutrients from both plant and animal materials, no generalizations about
the anatomical features of all omnivores can realistically be made.The variety of different
animals that are classified as omnivores can be placed into further sub-categories depending
on their feeding behaviors.
Frugivores include maned wolves and orangutans; insectivores include swallows and pink fairy
armadillos; granivores include large ground finches and mice.
All of these animals are omnivores, yet still fall into special niches in terms of feeding
behavior and preferred foods.
Being omnivores gives these animals more food security in stressful times or makes possible
living in less consistent environments.
== Etymology and definitions == The word omnivore derives from the Latin omnis
(all), and vora, from vorare, (to eat or devour), having been coined by the French and later
adopted by the English in the 1800s.
Traditionally the definition for omnivory was entirely behavioral by means of simply
"including both animal and vegetable tissue in the diet."
In more recent times, with the advent of advanced technological capabilities in fields like
gastroenterology, biologists have formulated a standardized variation of omnivore used
for labeling a species' actual ability to obtain energy and nutrients from materials.
This has subsequently conditioned two context specific definitions.
Behavioral: This definition is used to specify if a species or individual is actively consuming
both plant and animal materials.
(e.g. "vegans do not participate in the omnivore based diet.")
Physiological: This definition is often used in academia to specify species that have the
capability to obtain energy and nutrients from both plant and animal matter.
(e.g. "humans are omnivores due to their capability to obtain energy and nutrients from both plant
and animal materials.")The taxonomic utility of omnivore's traditional and behavioral definition
is limited, since the diet, behavior, and phylogeny of one omnivorous species might
be very different from that of another: for instance, an omnivorous pig digging for roots
and scavenging for fruit and carrion is taxonomically and ecologically quite distinct from an omnivorous
chameleon that eats leaves and insects.
The term "omnivory" is also not always comprehensive because it does not deal with mineral foods
such as salt licks and the consumption of plant and animal material for medical purposes
which would not otherwise be consumed (i.e. zoopharmacognosy) within non-omnivores.
=== Classification, contradictions and difficulties ===
Though Carnivora is a taxon for species classification, no such equivalent exists for omnivores, as
omnivores are widespread across multiple taxonomic clades.
The Carnivora order does not include all carnivorous species, and not all species within the Carnivora
taxon are carnivorous.
It is common to find physiological carnivores consuming materials from plants or physiological
herbivores consuming material from animals, e.g. felines eating grass and deer eating
birds.
From a behavioral aspect, this would make them omnivores, but from the physiological
standpoint, this may be due to zoopharmacognosy.
Physiologically, animals must be able to obtain both energy and nutrients from plant and animal
materials to be considered omnivorous.
Thus, such animals are still able to be classified as carnivores and herbivores when they are
just obtaining nutrients from materials originating from sources that do not seemingly complement
their classification.
For instance, it is well documented that animals such as giraffes, camels, and cattle will
gnaw on bones, preferably dry bones, for particular minerals and nutrients.
Felines, which are usually regarded as obligate carnivores, occasionally eat grass to regurgitate
indigestibles (e.g. hair, bones), aid with hemoglobin production, and as a laxative.Occasionally,
it is found that animals historically classified as carnivorous may deliberately eat plant
material.
For example, in 2013, it was considered that American alligators (Alligator mississippiensis)
may be physiologically omnivorous once investigations had been conducted on why they occasionally
eat fruits.
It was suggested that alligators probably ate fruits both accidentally but also deliberately."Life-history
omnivores" is a specialized classification given to organisms that change their eating
habits during their life cycle.
Some species, such as grazing waterfowl like geese, are known to eat mainly animal tissue
at one stage of their lives, but plant matter at another.
The same is true for many insects, such as beetles in the family Meloidae, which begin
by eating animal tissue as larvae, but change to eating plant matter after they mature.
Likewise, many mosquito species in early life eat plants or assorted detritus, but as they
mature, males continue to eat plant matter and nectar whereas the females (such as those
of Anopheles, Aedes and Culex) also eat blood to reproduce effectively.
== Omnivorous species ==
=== General ===
Although cases exist of herbivores eating meat and carnivores eating plant matter, the
classification "omnivore" refers to the adaptations and main food source of the species in general,
so these exceptions do not make either individual animals or the species as a whole omnivorous.
For the concept of "omnivore" to be regarded as a scientific classification, some clear
set of measurable and relevant criteria would need to be considered to differentiate between
an "omnivore" and other categories, e.g. faunivore, folivore, and scavenger.
Some researchers argue that evolution of any species from herbivory to carnivory or carnivory
to herbivory would be rare except via an intermediate stage of omnivory.
=== Omnivorous mammals === Various mammals are omnivorous in the wild,
such as species of pigs, badgers, bears, coatis, civets, hedgehogs, opossums, skunks, sloths,
squirrels, raccoons, chipmunks, mice, and rats.
Hominidae, including humans and chimpanzees, are also omnivores.
Most bear species are omnivores, but individual diets can range from almost exclusively herbivorous
to almost exclusively carnivorous, depending on what food sources are available locally
and seasonally.
Polar bears are classified as carnivores, both taxonomically (they are in the order
Carnivora), and behaviorally (they subsist on a largely carnivorous diet).
Depending on the species of bear, there is generally a preference for one class of food,
as plants and animals are digested differently.
Wolf subspecies (including wolves, dogs, dingoes, and coyotes) eat some plant matter, but they
have a general preference and are evolutionarily geared towards meat.
Also, the maned wolf is a canid whose diet is naturally 50% plant matter.
While most mammals may display "omnivorous" behavior patterns depending on conditions
of supply, culture, season and so on, they will generally prefer a particular class of
food, to which their digestive processes are adapted.
Like most arboreal species, most squirrels are primarily granivores, subsisting on nuts
and seeds.
But like virtually all mammals, squirrels avidly consume some animal food when it becomes
available.
For example, the American eastern gray squirrel has been introduced by humans to parts of
Britain, continental Europe and South Africa.
Where it flourishes, its effect on populations of nesting birds is often serious, largely
because of consumption of eggs and nestlings.
=== Other species === Various birds are omnivorous, with diets varying
from berries and nectar to insects, worms, fish, and small rodents.
Examples include cassowaries, chickens, crows and related corvids, keas, rallidae, and rheas.
In addition, some lizards, turtles, fish (such as piranhas and catfish), and invertebrates
are also omnivorous.
Quite often, mainly herbivorous creatures will eagerly eat small quantities of animal
food when it becomes available.
Although this is trivial most of the time, omnivorous or herbivorous birds, such as sparrows,
often will feed their chicks insects while food is most needed for growth.
On close inspection it appears that nectar-feeding birds such as sunbirds rely on the ants and
other insects that they find in flowers, not for a richer supply of protein, but for essential
nutrients such as cyanocobalamin that are absent from nectar.
Similarly, monkeys of many species eat maggoty fruit, sometimes in clear preference to sound
fruit.
When to refer to such animals as omnivorous, or otherwise, is a question of context and
emphasis, rather than of definition.
== See also